Evolution is not changes in gene frequencies. Genes are part of a network of developmental causes that lead to the manifestation of traits that have general properties in common across individuals while retaining individual differences. Evolution is change in the frequencies of alternative developmental causes that yield variations in developmental trajectories (a phrase that is more cumbersome than ‘‘changes in gene frequencies’’ but nevertheless more correct).
Source: Michel, GF and Tyler, AN. Developing human nature:“Development to” versus “Development from?”. Developmental Psychobiology (2007) vol. 49 (8) pp. 788-799 DOI: 10.1002/dev.20261